nonparametric bootstrap files contain only zeros

[Maria Paula Rodriguez]

Hi Ryan,

I'm new in dadi so maybe I'm not following the code correctly, but I couldn't find any conversation to address this issue (maybe the "Nonparametric bootstrapping" could help, but I don't think my SNPs have the same name, see below).

I am trying to generate non-parametric bootstrapping of my data, but all generated files have only zeros. I couldn't attached files, so please find how two bootstrapped FS look like at the end of this message (I inspected several of them and they all look the same).

I have used 10e7 and 2e6 as size chunks, but with both I am getting the same files with only 0s. I generated the SFS of my data with easySFS. I conducted the down-projection by maximizing the number of SNPs per population, where I kept 138 and 180 samples. This is the complete code I am using for the non-parametric bootstrapping:

fs = dadi.Spectrum.from_file("./downstreamM-upstreamM.sfs")
min_value = fs.min()
print(min_value) # lowest value is 7.63775735436711e-24

# Creating and saving the dictionary
vcf = "./Data.vcf"
vcfDict = dadi.Misc.make_data_dict_vcf(vcf, './popAssign_OneDownOneUp.txt')
import os
if not os.path.exists('./data_dictionaries'):
  os.makedirs('./data_dictionaries')

pick = open('./ DownMig.UppMig.vcf.dict', 'wb')
pickle.dump(vcfDict, pick, 2)

# Uploading the dictionary
vcfDict =  pickle.load(open('./DownMig.UppMig.vcf.dict','rb'))

# Creating and saving FS for bootstraped genomes
Nboot, chunk_size = 100, 2e6
chunks = dadi.Misc.fragment_data_dict(vcfDict, chunk_size)
pop_ids, ns = ["downstreamM", "upstreamM"] , [138, 180]
boots = dadi.Misc.bootstraps_from_dd_chunks(chunks, Nboot, pop_ids, ns)

# Save the bootstraps
for i in range(len(boots)):  boots[i].to_file('./boots/UppM.DownMig.trial2.subset.boot_{0}.fs'.format(str(i)))


I am also attaching the 1D and 2D spectra of my populations, because I don't know if the problem may be due to the FS. However, the number of snp's is similar to the example 1D spectra on the dadi website. I also attach a photo with a portion of the data dictionary generated by easySFS (datadict) , which shows that the snp's id's are not identical (NC_048565.1-27105-6 vs NC_048565.1-144286-11). I am assuming that this file follows the same structure than the dictionaries created with dadi.Misc.make_data_dict_vcf (I may be wrong). I've used the former function to create it, but when I print it to assess it content, it seems that I can only see binary values. Nevertheless, I am using the dictionary created with dadi for the bootstrapping.

I haven't created all my demographic models because I first want to know if my FS input is OK and what is creating the problem with the bootstrap FS before moving forward. I would really appreciate any suggestion or clarification about this.

Thank you in advance

# Bootstap file 1
139 181 unfolded "downstreamM" "upstreamM"
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[Ryan Gutenkunst]

Hello Maria,

The issue is with the VCF file and SNP designations. The fragment_data_dict function expects keys to be of the form "chromosome_position[.additional_info]” which yours aren’t https://dadi.readthedocs.io/en/latest/api/dadi/Misc.html#dadi.Misc.fragment_data_dict .

I’m not sure how to read the SNP ideas you do have, but you could write a simple function to parse them into the format dadi expects.

Best,

Ryan

<1DSpectra.PNG>

<2DSpectra.PNG>

<datadict.PNG>

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[Maria Paula Rodriguez]

Hi Ryan,

Thanks for your prompt response. 

I modified the SNP's names in the vcf, run easySFS to generate the FS using the new vcf and then created the dictionary again. However, the problem persists. I also avoided having SNP's names that start with 0, it seems that python doesn't like it. However, I don't know what else to try. Here is my new code (I'm showing the structure of the dictionary and it seems to be in the expected format for the function):

# In case it's helpful, this is the command I'm using in easySFS:

./easySFS.py -i  ./DownMigrant.UppeMig.SNPsdadi2.vcf   -p  ./popAssign_OneDownOneUp.txt  -a --proj=138,180

# New FS

fs = dadi.Spectrum.from_file("./downstreamM-upstreamM.sfs")

print(fs)

[[-- 50364.40973840228 19506.52545925824 ... -- -- --] [37822.78653369307 21542.67582831754 12012.46117673672 ... -- -- --] [9776.239861528618 7356.083052229609 5033.494801393011 ... -- -- --] ... [8.88972787591124e-22 8.182693911379928e-20 3.620665138082356e-18 ... -- -- --] [1.240025583782837e-22 1.141390025356761e-20 5.050344781452131e-19 ... -- -- --] [7.63775735436711e-24 7.030211704540504e-22 3.110672857641352e-20 ... -- -- --]]

# the 2D plot looks the same as before

# New vcf

vcf = "./DownMigrant.UppeMig.SNPsdadi2.vcf"

vcfDict = dadi.Misc.make_data_dict_vcf(vcf, './popAssign_OneDownOneUp.txt')

# Does the dictionary have the expected format? (chrom_pos): yes

first_four = dict(islice(vcfDict.items(), 4))

print(first_four)

{'chr048565_20048': {'segregating': ('G', 'A'), 'context': '-G-', 'outgroup_allele': '-', 'outgroup_context': '---', 'calls': {'downstreamM': (0, 194), 'upstreamM': (0, 296)}}, 'chr048565_20049': {'segregating': ('T', 'A'), 'context': '-T-', 'outgroup_allele': '-', 'outgroup_context': '---', 'calls': {'downstreamM': (0, 194), 'upstreamM': (0, 296)}}, 'chr048565_20087': {'segregating': ('C', 'A'), 'context': '-C-', 'outgroup_allele': '-', 'outgroup_context': '---', 'calls': {'downstreamM': (0, 194), 'upstreamM': (0, 296)}}, 'chr048565_20111': {'segregating': ('T', 'G'), 'context': '-T-', 'outgroup_allele': '-', 'outgroup_context': '---', 'calls': {'downstreamM': (0, 194), 'upstreamM': (0, 296)}}}

# Bootstrap

Nboot, chunk_size = 100, 2e7
chunks = dadi.Misc.fragment_data_dict(vcfDict, chunk_size)

pop_ids, ns = ["downstreamM", "upstreamM"] , [138, 180] # I'm using the same samples as the downprojection in easySFS

boots = dadi.Misc.bootstraps_from_dd_chunks(chunks, Nboot, pop_ids, ns)

# content of boots fs: all have 0's and a single 1 as the last entry (I also opened the files) 

print(boots[13])

[[-- 0.0 0.0 ... 0.0 0.0 0.0] [0.0 0.0 0.0 ... 0.0 0.0 0.0] [0.0 0.0 0.0 ... 0.0 0.0 0.0] ... [0.0 0.0 0.0 ... 0.0 0.0 0.0] [0.0 0.0 0.0 ... 0.0 0.0 0.0] [0.0 0.0 0.0 ... 0.0 0.0 --]]

I'd appreciate any further guidance.

Thank you in advance

[Ryan Gutenkunst]

Hello Maria,

I think the issue now is that boostraps_from_dd_chunks defaults to polarized=True , but you don’t have outgroup information in your data dictionary. If you set polarized=False, I suspect it will work.

Best,

Ryan

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[Maria Paula Rodriguez]

Hi Ryan

It worked!! Thank you very much. 

Maria

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