HI Aaron,
So tuning does occur in the model itself automatically. However, I always check to see if the model is overfit and often find that it is. To do this I compare the cross-validation statistics to the training statistics. If the values are very different, I go back to the learning rate and tree complexity, varying these using the parameter explore option in VisTrails to optimize them to come up with the highest assessment metrics that are also close together for the training and CV.
I hope that helps. There's a good explanation in the Elith paper on BRT that I believe is referenced in the documentation.
Catherine