Afingerprint is an impression left by the friction ridges of a human finger. The recovery of partial fingerprints from a crime scene is an important method of forensic science. Moisture and grease on a finger result in fingerprints on surfaces such as glass or metal. Deliberate impressions of entire fingerprints can be obtained by ink or other substances transferred from the peaks of friction ridges on the skin to a smooth surface such as paper. Fingerprint records normally contain impressions from the pad on the last joint of fingers and thumbs, though fingerprint cards also typically record portions of lower joint areas of the fingers.
Human fingerprints are detailed, nearly unique, difficult to alter, and durable over the life of an individual, making them suitable as long-term markers of human identity. They may be employed by police or other authorities to identify individuals who wish to conceal their identity, or to identify people who are incapacitated or deceased and thus unable to identify themselves, as in the aftermath of a natural disaster.
Their use as evidence has been challenged by academics, judges and the media. There are no uniform standards for point-counting methods, and academics have argued that the error rate in matching fingerprints has not been adequately studied and that fingerprint evidence has no secure statistical foundation.[1] Research has been conducted into whether experts can objectively focus on feature information in fingerprints without being misled by extraneous information, such as context.[2]
Fingerprints are impressions left on surfaces by the friction ridges on the finger of a human.[3] The matching of two fingerprints is among the most widely used and most reliable biometric techniques. Fingerprint matching considers only the obvious features of a fingerprint.[4]
The composition of fingerprints consists of water (95%-99%), as well as organic and inorganic constituents.[5] The organic component is made up of amino acids, proteins, glucose, lactase, urea, pyruvate, fatty acids and sterols.[5] Inorganic ions such as chloride, sodium, potassium and iron are also present.[5] Other contaminants such as oils found in cosmetics, drugs and their metabolites and food residues may be found in fingerprint residues.[6]
A friction ridge is a raised portion of the epidermis on the digits (fingers and toes), the palm of the hand or the sole of the foot, consisting of one or more connected ridge units of friction ridge skin.[citation needed] These are sometimes known as "epidermal ridges" which are caused by the underlying interface between the dermal papillae of the dermis and the interpapillary (rete) pegs of the epidermis. These unique features are formed at around the 15th week of fetal development and remain until after death, when decomposition begins.[7] During the development of the fetus, around the 13th week of a pregnancy, ledge-like formation is formed at the bottom of the epidermis beside the dermis.[7] The cells along these ledges begin to rapidly proliferate.[7] This rapid proliferation forms primary and secondary ridges.[7] Both the primary and secondary ridges act as a template for the outer layer of the skin to form the friction ridges seen on the surface of the skin.[7]
These epidermal ridges serve to amplify vibrations triggered, for example, when fingertips brush across an uneven surface, better transmitting the signals to sensory nerves involved in fine texture perception.[8] These ridges may also assist in gripping rough surfaces and may improve surface contact in wet conditions.[9]
Consensus within the scientific community suggests that the dermatoglyphic patterns on fingertips are hereditary.[10] The fingerprint patterns between monozygotic twins have been shown to be very similar (though not identical), whereas dizygotic twins have considerably less similarity.[10] Significant heritability has been identified for 12 dermatoglyphic characteristics.[11] Current models of dermatoglyphic trait inheritance suggest Mendelian transmission with additional effects from either additive or dominant major genes.[12]
Whereas genes determine the general characteristics of patterns and their type, the presence of environmental factors result in the slight differentiation of each fingerprint. However, the relative influences of genetic and environmental effects on fingerprint patterns are generally unclear. One study has suggested that roughly 5% of the total variability is due to small environmental effects, although this was only performed using total ridge count as a metric.[10] Several models of finger ridge formation mechanisms that lead to the vast diversity of fingerprints have been proposed. One model suggests that a buckling instability in the basal cell layer of the fetal epidermis is responsible for developing epidermal ridges.[13] Additionally, blood vessels and nerves may also serve a role in the formation of ridge configurations.[14] Another model indicates that changes in amniotic fluid surrounding each developing finger within the uterus cause corresponding cells on each fingerprint to grow in different microenvironments.[15] For a given individual, these various factors affect each finger differently, preventing two fingerprints from being identical while still retaining similar patterns.
It is important to note that the determination of fingerprint inheritance is made difficult by the vast diversity of phenotypes. Classification of a specific pattern is often subjective (lack of consensus on the most appropriate characteristic to measure quantitatively) which complicates analysis of dermatoglyphic patterns. Several modes of inheritance have been suggested and observed for various fingerprint patterns. Total fingerprint ridge count, a commonly used metric of fingerprint pattern size, has been suggested to have a polygenic mode of inheritance and is influenced by multiple additive genes.[10] This hypothesis has been challenged by other research, however, which indicates that ridge counts on individual fingers are genetically independent and lack evidence to support the existence of additive genes influencing pattern formation.[16] Another mode of fingerprint pattern inheritance suggests that the arch pattern on the thumb and on other fingers are inherited as an autosomal dominant trait.[17] Further research on the arch pattern has suggested that a major gene or multifactorial inheritance is responsible for arch pattern heritability.[18] A separate model for the development of the whorl pattern indicates that a single gene or group of linked genes contributes to its inheritance.[19] Furthermore, inheritance of the whorl pattern does not appear to be symmetric in that the pattern is seemingly randomly distributed among the ten fingers of a given individual.[19] In general, comparison of fingerprint patterns between left and right hands suggests an asymmetry in the effects of genes on fingerprint patterns, although this observation requires further analysis.[20]
In addition to proposed models of inheritance, specific genes have been implicated as factors in fingertip pattern formation (their exact mechanism of influencing patterns is still under research). Multivariate linkage analysis of finger ridge counts on individual fingers revealed linkage to chromosome 5q14.1 specifically for the ring, index, and middle fingers.[21] In mice, variants in the gene EVI1 were correlated with dermatoglyphic patterns.[22] EVI1 expression in humans does not directly influence fingerprint patterns but does affect limb and digit formation which in turn may play a role in influencing fingerprint patterns.[22] Genome-wide association studies found single nucleotide polymorphisms within the gene ADAMTS9-AS2 on 3p14.1, which appeared to have an influence on the whorl pattern on all digits.[23] This gene encodes antisense RNA which may inhibit ADAMTS9, which is expressed in the skin. A model of how genetic variants of ADAMTS9-AS2 directly influence whorl development has not yet been proposed.[23]
In February 2023, a study identified WNT, BMP and EDAR as signaling pathways regulating the formation of primary ridges on fingerprints, with the first two having an opposite relationship established by a Turing reaction-diffusion system.[24][25][26]
Before computerization, manual filing systems were used in large fingerprint repositories.[27] A fingerprint classification system groups fingerprints according to their characteristics and therefore helps in the matching of a fingerprint against a large database of fingerprints. A query fingerprint that needs to be matched can therefore be compared with a subset of fingerprints in an existing database.[4] Early classification systems were based on the general ridge patterns, including the presence or absence of circular patterns, of several or all fingers. This allowed the filing and retrieval of paper records in large collections based on friction ridge patterns alone. The most popular systems used the pattern class of each finger to form a numeric key to assist lookup in a filing system. Fingerprint classification systems included the Roscher System, the Juan Vucetich System and the Henry Classification System. The Roscher System was developed in Germany and implemented in both Germany and Japan. The Vucetich System was developed in Argentina and implemented throughout South America. The Henry Classification System was developed in India and implemented in most English-speaking countries.[27]
The numbers assigned to each print are based on whether or not they are whorls. A whorl in the first fraction is given a 16, the second an 8, the third a 4, the fourth a 2, and 0 to the last fraction. Arches and loops are assigned values of 0. Lastly, the numbers in the numerator and denominator are added up, using the scheme:
Fingerprint identification, known as dactyloscopy,[30] ridgeology,[31] or hand print identification, is the process of comparing two instances of friction ridge skin impressions (see minutiae), from human fingers or toes, or even the palm of the hand or sole of the foot, to determine whether these impressions could have come from the same individual. The flexibility and the randomized formation of the friction ridges on skin means that no two finger or palm prints are ever exactly alike in every detail; even two impressions recorded immediately after each other from the same hand may be slightly different.[30] Fingerprint identification, also referred to as individualization, involves an expert, or an expert computer system operating under threshold scoring rules, determining whether two friction ridge impressions are likely to have originated from the same finger or palm (or toe or sole).
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