Hi Jacob,
Thanks for the question. This problem could arise for a variety of reasons. First off, it certainly seems like the random effect variance is unidentifiable in this case given the massive credible interval. This could be a result of the data for this species not being able to estimate all parameters in this model (e.g., if there are a small number of detections, the detections are highly clustered in a small set of the sites). I would not recommend simply trying to set a stricter prior on sigma.sq.psi, as this suggests to me there is not enough data to identify the problem. I would encourage you to take a look at the traceplots of the random effect estimates (i.e., both the variance in "sigma.sq.psi.samples" as well as the actual estimated random effect intercepts stored in beta.star.samples). Looking at those values may help reveal the root cause of what's causing the problems with estimating the random effect. In your simpler models that you say converge, I would also look at the magnitude of the credible intervals for the variance. Is it similarly large, or are the values more reasonable? If they are more reasonable, this likely indicates the full model is too complex and the data for the species can't estimate the parameters. If it is similarly large, the model still may just not have much information to estimate the random effect. Regardless, I'd encourage you to dig into the actual locations of where the detections for this species are and how that might be contributing to the challenge in estimating the RE variances.
Hope that helps,
Jeff