Hi there,
Thanks for your message! I have tried to reproduce the error but have not been able to -- MCMCtree runs without problems when using PAML v4.10.7. Which PAML version are you using? Also, make sure that the alignment is correct as it seems you have a lot of empty sequences.
All the best,
S.
P.S.: I guess that you just included the input files for the first beta point, but make sure that you run MCMCtree for the rest of the beta points as the tutorial specifies too!