Autocrat does not have its own IP address because it makes use of Google's network of IP addresses.(This add-on is built on the Google Platform, using app script and Google mail servers to complete the mailing actions.)
When Autocrat pushes out an email from your spreadsheet it sends the email on behalf of your account but these are emails are run by a script and will use whatever Google server is available at the time to send it out. The server can change from email to email but it is always a Google server. My understanding is that there is not a set list of Google IP addresses' you can whitelist.
You should check this with Google Support/ Mimecast Support but I would guess that means all Gmail addresses would need to be whitelisted which I understand is an issue in terms that it leaves you open to all Gmail spoof emails.
Apologies that I do not have specific steps to resolve this issue, many times email filtering software does not play nice with Autocrat or other third-party add-ons that use app script triggers to bulk email. I hope with some of this information you and your IT department can brainstorm other solutions that could work with your domain's email security requirements.
Best of luck in getting this issue resolved. Please keep us posted on your progress.