Hello,
First off, your live website is absolutely fantastic and so far the most useful resource I've come across for explaining microbial ecology work!
In the section RDA> Pre-analysis > step 3: it is recommended to standardize your explanatory data if they are not dimensionally homogeneous.
I've looked up this topic in the past and it seems that doing so depends on the method of your analysis. As Jari Oksanen explains (
https://stat.ethz.ch/pipermail/r-sig-ecology/2010-April/001196.html) this is not necessary in vegan. Might be worth updating to clarify that portion.