Myidream,
This is a general problem dating back to the origins of LANDIS. It is always possible to push the model beyond its intended capacity. And simulating 400 million ha for 300 years using Century is well beyond the reach of today's common computing platforms.
A few suggestions:
* Choose a smaller area. Or choose a model without spatial interactions. Are spatial interactions necessary and useful across 400 M ha? How important is seed dispersal, in particular, to your answering your questions?/
* Choose a larger resolution. You didn't mention your resolution but the CPU cost of seed dispersal increases exponentially as cell size decreases linearly (it is a Big-O-squared type algorithm).
* Use an extension that carries less information. Century Succession uses the MOST memory of any extension as it is tracking many C and N pools. If tracking N is not critical, there are other C-centric succession to choose from, including 'Forest Carbon' (available now) and 'PnET' (available soon).
Cheers,
Rob