Also relevant is
20. Lantana strigocamara R.W. Sanders, Sida 22:392. 2006. Type: U.S.A. florida. Dade Co.: Coral Gables, 23 Sep 1981, Sanders 1450 (holotyPe: FTG!; isotyPe: NY!).
Lantana mutabilis Lippold ex Otto & A. Dietr., Allg. Gartenzeitung 10:314. 1842. nom. illeg. tyPe: Unknown.
Shrubs erect or rounded, open; stems 0.3–3 m; branches ascending, several to numerous; twigs, peduncles and often petioles thinly to moderately strigose, setose, or pilose, the hairs 0.1–1.2(–1.5) mm, the longest mostly 0.5–1 mm. Leaf-blades ovate to broadly ovate, (2–)5–10 cm long, the length 1–1.7 × width, usually not nigrescent, papery, pinninerved; base rounded, truncate, or cordate, shortly and narrowly cuneate onto petiole at very base; apex usually acuminate; marginal teeth 15–40 per side, rounded to acute, often appressed, sometimes spreading at tip, with sinuses 0.5–1.5 mm deep; adaxial surface usually dull, antrorsely strigose or strigose-setose, the hairs occurring on veins and intervening tissue (sometimes just center of areoles), 0.2–1.2 (longest mostly 0.5–0.8) mm, 1–12/sq. mm, not noticeably vitreous-pustulate, the circular bases of the strigae ca. 0.1–0.2 mm in diam.; abaxial surface slightly lighter or duller green than adaxial surface, antrorsely strigose-scabrous, with the strigae scattered to moderately dense on veins and veinlets, 0.1–0.6 (longest ones usually 0.4–0.6) mm (sometimes accompanied by scattered short [mostly ≤ 0.3 mm] erect filiform hairs along major veins), 4–20/sq. mm. Inflorescences remaining hemispheric; peduncles 0.5–1.2 × leaf length. Proximal bracts narrowly triangular, linear-lanceolate, or linear-oblong (including those with slight constriction in proximal third; occasionally 1 or 2 outermost bracts subfoliar or narrowly spatulate), (3–)5–8(–10) mm long, 0.8–1.5 (rarely an occasional subfoliar bract up to 2) mm wide, widest at or just above the base, with 3 veins from the base, appressed or spreading, deciduous after flowering; apex acute to attenuate; indument strigose or strigillose, hardly ciliate, the longest hairs 0.2–0.6 mm. Corolla opening yellow or creamy white (rarely pure white) with yellow throat, aging to cream, dark yellow, orange, or red (rarely remaining white), often infused with pink or purple; corolla tube 7–12 mm.
Distribution and habitat.—Of cultivated origin; cultivated worldwide and escaped pantropically (especially southern United States, Caribbean Basin, India, Sri Lanka, Southeast Asia, and tropical Africa); open woodland, thickets, disturbance openings and man-made grassland, calcareous or sandy soils; 0–2000 m. Sanders (2006) discussed Lantana strigocamara at length, and Sanders (1987a) illustrated the species (as “camara”).
In view of above, I am reproducing keys to all species here:
Key to species of lantana sect. lantana
Before attempting to use the identification key or descriptions, see caveats above.
1. Abaxial leaf-surface antrorsely strigose-scabrous to nearly glabrous, consisting only of or strongly dominated by strigae.
2. Proximal bracts 2–8 mm wide, with 5–7 veins from the base.
3. Capitula not elongating by prolonged initiation of additional flowers, remaining hemispheric; bracts ± obtuse, acute and rounded at very tip, or briefly acuminate, appressed to spreading, appearing to form an involucre; cilia, if present on bracts, usually no more than 0.5 mm; corollas yellow or orange aging reddish (rarely intensely reddish purple) ___________ 13. L. cujabensis
3. Capitula elongating by prolonged initiation of additional flowers, becoming cylindric; bracts acuminate with a prolonged tip, spreading, causing the capitula to resemble spikes of Carex lupulina; cilia usually well developed on bracts, mostly 0.5–1 mm; corollas pink to purple or white aging purplish (rarely yellow to red-orange) _______ 19. L. paraensis
2. Proximal bracts 0.5–1.5 mm wide (to 3 mm in L. ovatifolia and L. kingii, otherwise rarely one or two bracts in outermost series spatulate-subfoliaceous to about 2 mm wide), with 3 veins (rarely 4 or 5 in L. splendens) from the base.
4. Leaf-blades ovate-elliptic to lanceolate-elliptic or trullate, averaging 1.7–2.5(–3) times longer than wide, the base attenuately tapering to petiole from middle or just below middle, or less commonly abruptly contracted and broadly cuneate onto petiole; blades triplinerved; herbage usually nigrescent (except in L. kingii and L. splendens); upper leaf-surface usually lustrous.
5. Proximal bracts (excluding one or two subfoliar outermost ones or those of gall-transformed heads) widest near or just below middle or in distal half (sometimes so in L. splendens, see below), persistent in fruit.
6. Leaf-blades bright, dark, or dull green abaxially, nigrescent, ovate-elliptic, induplicate or having halves incurved at maturity; larger strigae of the abaxial leaf-surface 0.5–1 mm; twigs setulose with spreading hairs about 0.5–1.5 mm; proximal bracts ca. 3–5 mm, elliptic-lanceolate ________________________________________9. L. depressa
6. Leaf-blades distinctly whitish or pale green below (though not glaucous), not nigrescent, usually ovate or ovate-triangular, rarely (especially if less than 2 cm long) obovate or ovate-elliptic, ± flat, not having halves incurved at maturity; larger strigae of the abaxial leaf-surface 0.1–0.4(–0.6) mm; twigs glabrescent with antrorse hairs 0.3–0.7 mm; proximal bracts ca. 6–10 mm long, spatulate or oblanceolate __________________________10. L. kingii
5. Proximal bracts (excluding one or two subfoliar outermost ones or those of gall-transformed heads) widest at or just above base, deciduous after flowering.
7. Corollas white aging bluish to pink, or pink aging light purple, or with purple intermixed with creamy yellow to orange; hairs of abaxial leaf-surface moderately dense, ca. 15–60/mm2 (under 10X magnification those on the higher order veins may be so small as to appear to be papillae); abaxial surface of leaf-blade not noticeably whitish-green; stems often with abundant, stout, recurved prickles ___________________________________12. L. nivea
7. Corollas yellow to reddish orange without any pink or purple mixed in; hairs of abaxial leaf-surface moderately to very sparse, ca. 0–12/mm2 (under 10X magnification those on the higher order veins do not appear to be papillae); abaxial surface of leaf-blade whitish-green (but not glaucous); stems usually lacking stout, recurved prickles.
8. Leaf-blades mostly 1–5(–7) cm long, on adaxial surface of mature and older leaves the circular bases of strigae 0.3–0.5 or more mm in diam., conspicuously vitreous-pustulate, often nearly filling whole areole; corolla tubes ca. 5–8 mm long; proximal bracts mostly oblong-lanceolate (outermost 1 or 2 oblong-obovate or oblanceolate), 2–4(–5) mm long; virgate or divaricately branched shrubs ________________________7. L. splendens
8. Leaf-blades mostly 5–15 cm long, on adaxial surface the circular bases of strigae usually 0.2 mm or less in diam. (not exceeding 0.3 mm.), usually not conspicuously vitreous-pustulate, not filling whole areole; corolla tubes 7–12 mm long; proximal bracts narrowly lanceolate to linear-triangular, 4–10 mm long; rounded, lax, or subscandent shrubs.
9. Leaf-blades glabrescent, smooth and subsucculent or coriaceous; hairs of adaxial leaf-surface geniculately bent at very base, flaccid, strongly appressed to surface and often deciduous; peduncles a third or less as long as leaves ______8. L. hodgei
9. Leaf-blades strigose or scabrous, papery to subcoriaceous; hairs of adaxial leaf-surfaces geniculately bent about 1⁄4 of length above base and held above surface; peduncles about equaling to half as long as leaves __ 6. L. scabrida
4. Leaf-blades ovate to broadly ovate, averaging 1–1.7 times longer than wide, the base rounded, truncate, or cordate, usually briefly and narrowly cuneate onto petiole at very base of blade; blades pinninerved; herbage usually not nigrescent (if so, then only the young, expanding leaves); upper leaf-surface lustrous or not.
10. Adaxial leaf-surface dull, not vitreous-pustulate, the bases of the strigae only about 0.1–0.2 mm in diam.; bracts deciduous after anthesis, the proximal series lanceolate-triangular or lanceolate-linear (occasionally an outermost one or two spatulate-subfoliar); corollas opening yellow, cream, or white aging yellow to red-orange, purplish, orange plus purple, or white with yellow throat _____ 20. L. strigocamara
10. Adaxial leaf-surface lustrous, vitreous-pustulate or pustulate-scabridulous, the bases of the strigae (at least on the older leaves) about 0.3–0.5 mm or more in diam.; bracts persistent into fruit, the proximal series typically spatulate to elliptic-oblong; corollas yellowish to orange or red-orange.
11. Hairs on adaxial leaf-surface mostly 0.1–0.4 mm, appressed, often deciduous leaving only pustulate bases; leaf-blades abaxially distinctly whitish or pale green (seemingly but not actually glaucous); leaf-teeth sinuses 1–2.5 mm deep; stems upright; corollas opening yellow aging orange or red-orange ___________________10. L. kingii
11. Hairs on adaxial leaf-surface 0.2–1 mm, ascending, longer ones sometimes deciduous; leaf-blades abaxially bright, dark or dull green; leaf-teeth sinuses 0.7–1.5 mm deep; stems trailing or decumbent; corollas opening and ± remaining yellow ______________________________________________________________ 11. L. ovatifolia
1. Abaxial leaf-surface not exclusively or dominantly antrorsely strigose-scabrous but setose, pilose, velutinous, puberulent, pannose, viscid, or glabrescent, the vestiture varying from having hairs that are exclusively erect (± erect from basal insertion, spreading from surface of lamina or vein from which they arise, filiform or setaceous, gland-tipped or not, distally arching-curved or flexuously curled) without strigae present to having a codominant mixture of erect hairs and strigae (occurs in hybrids and some species of hybrid origin).
12. Capitula elongating, becoming cylindric; corollas usually pink to deep reddish purple (sometimes pale yellow in throat only), occasionally white becoming infused with purple (rarely yellow to orange red in L. paraensis).
13. Corolla tubes briefly or not exerted beyond bracts, 2–4 mm; capitula elongating by expansion of nodes between flowers/fruits; bracts often deciduous after flowering, but if persisting, then usually rapidly becoming reflexed from base, without cilia, abaxial hairs 0.3–0.5 mm, apex acute to rounded, sometimes abruptly acuminate or briefly attenuate _______15. L. micrantha
13. Corolla tubes exerted well beyond bracts, (5–)7–12 mm; capitula elongating by prolonged initiation of flowers; bracts strongly persisting and spreading in fruit, with cilia or also abaxial hairs (0.5–)0.7–1.5 mm, apex acuminate with prolonged tip.
14. Hairs of leaf-surfaces 30–150/mm2, longest ones 1–1.5 mm or more; twigs and peduncles densely stipitate glandular with scattered setae _________ 14. L. viscosa
14. Hairs of leaf-surfaces 3–20/mm2, longest ones 0.3–0.7 (rarely to 1.2) mm; twigs and peduncles glabrescent to thinly setulose, sometimes with scattered stipitate glands ______ 19. L. paraensis
12. Capitula not elongating, remaining hemispheric; corollas usually opening yellow or orange aging orange or redorange (sometimes opening creamy white or pure white and/or becoming infused with pink or purple in L. planaltensis, or rarely opening white and remaining so).
15. Leaves-blades distinctly triplinerved, usually nigrescent; adaxial leaf-surface often lustrous; abaxial surface often with strigae mixed with filiform hairs (can be covered over by the latter).
16. Leaf-blades mostly 2–5 cm long, mostly ovate-triangular or lanceolate-triangular with straight tapering sides from proximal 1⁄3 (sometimes contracted to abruptly acute or obtuse apex); adaxial surface usually noticeably lustrous and vitreous-pustulate, with the hair bases 0.3–0.5 mm diam., thinly scabrous, the strigae 2–10/mm2, about 0.3 mm or less giving the surface a sandpapery texture; abaxial surface with soft, straight hairs restricted to crevices between the leaf-surface and the midrib (or also secondary veins) _____________ 16. L. bahamensis
16. Leaf-blades mostly 5–10 cm long, ovate, lanceolate or ovate-elliptic with curved sides and usually acuminate apex; adaxial surface lustrous or not, usually not pustulate, with the hair bases less than 0.3 mm diam., strigosepilose, the hairs 20–80/ mm2 or more, of mixed length up to 0.7 mm; abaxial surface with even vestiture of soft, straight hairs on surfaces of midrib to higher order veins or also areole tissue _________________ 17. L. planaltensis
15. Leaves-blades pinninerved, not distinctly nigrescent (except in some L. urticoides); adaxial surface dull (except somewhat lustrous in some L. urticoides); abaxial surface lacking strigae (except in many interspecific hybrids), exclusively of filiform, glandular, or setiform hairs.
17. Leaves 1–1.5 times longer than wide, ± rotund, deltate or broadly ovate with conspicuous spreading acute teeth, the sinuses mostly 2–5 mm deep; adaxial leaf-surface, at least on older leaves noticeably vitreouspustulate, the bases of the strigae mostly 0.3–0.5 mm in diam; abaxial leaf-surface with long setaceous hairs restricted to the midrib and secondary veins, these gradually reduced in length from base of midrib (where 1.5–2 mm long) toward margin (on midrib and secondary veins to ca. 0.7 mm long), shortest hairs (0.2–0.5 mm long) restricted to veinlets and areoles; proximal bracts mostly 7–12 mm long, oblanceolate or spatulate, mostly 1.5–3 mm wide, widest in distal half or near middle, conspicuously persistent and reflexed in fruit __________18. L. urticoides
17. Leaves mostly 1.5–2.5 times longer than wide (if less than 1.5, then lacking character combination of lead 17’), ovate, oblong-deltate, elliptic, or lanceolate with rounded or appressed-acute teeth, the sinuses mostly 0.3–2 mm deep (to 3 mm in some L. hirsuta subsp. hirsuta); adaxial leaf-surface not vitreous-pustulate (except in some L. camara subsp. aculeata), the bases of the strigae 0.1–0.2(–0.3) mm in diam.; abaxial leaf-surface with hairs all about the same length (either long setaceous hairs only on veins or short soft hairs on veins and areoles; longer hairs scattered among shorter hairs on midrib in some L. horrida and some L. hirsuta, but not in pattern of L. urticoides); proximal bracts mostly 2–10 mm long, linear-lanceolate (rarely linear-spatulate), lanceolate- or elliptic-oblong, mostly 0.5–1.5 mm wide, widest in proximal third (if oblanceolate- or obovatespatulate and widest above middle, then mostly 2–6 mm long), deciduous or persistent (then sometimes reflexed) in fruit.
18. Hairs of abaxial leaf-surface setiform, ca. 0.7–1.5 mm, straight and erect, sinuous, or antrorsely arching, restricted mostly to midrib, secondary, and tertiary veins, without sparse understory of shorter (0.1–0.5 mm), softer filiform hairs; adaxial surface setose to villous dominated by antrorse setaceous hairs 1–2 mm between the secondary veins, sometimes these also accompanied by an understory of shorter hairs; young twigs (also petioles and peduncles) with spreading hairs (1.2–)1.5–2.5 mm.
19. Young twigs and peduncles usually viscid and sparsely setose, dominated by dense, conspicuous, stipitate glands to ca. 0.5 mm; proximal bracts mostly 4–6 mm long, oblong-elliptic or -lanceolate, covered with hairs ca. 1 mm and usually marginally ciliate with hairs (1–)1.5–2 mm ______________ 5. L. insularis
19. Young twigs and peduncles sparsely to moderately setose, stipitate glands lacking (except in interspecific hybrids); proximal bracts mostly 5–10 mm long, linear-lanceolate or linear-spatulate, covered with hairs 0.3–1 mm, marginally ciliate with hairs 0.8–1(–1.5) mm or these lacking _____________________4. L. hirsuta
18. Hairs of abaxial leaf-surface weak and filiform, 0.1–0.5 mm (sometimes in L. horrida, scattered arching hairs on midrib to 1 mm among shorter hairs), spreading to curled, usually occurring on all vein orders including veinlets and areoles, occasionally deciduous and persisting only in crevices between major veins and leaf-surface; adaxial surface antrorsely strigillose to strigose villous with a covering of hairs of mixed length, these mostly 0.1–0.9 mm; young twigs (also petioles and peduncles) with spreading to appressed hairs only 0.1–1 mm (to 1.5 mm in some L. horrida subsp. tilifolia).
20. Adaxial leaf-surface with a canopy of hairs between secondary veins only 0.2–0.5 mm (in L. camara subsp. aculeata sparsely scattered hairs to 0.7 mm may occur), usually in the form of strigae or stalked glands; peduncles typically about a third the length of to about equaling their subtending leaves (up to twice as long in L. camara subsp. glandulosissima) ________________________________________ 1. L. camara
20. Adaxial leaf-surface with a moderately dense canopy of hairs between secondary veins mostly 0.7– 0.8(–1) mm, in the form of arching or flexible setae, often with a well developed understory of shorter (≤ 0.5 mm) strigae, filiform hairs or stalked glands; peduncles typically longer to 3 times longer than subtending leaves.
21. Plants erect or trailing, laxly or openly branched, leaf-blades ovate to broadly elliptic to lanceolatedeltate, mostly 1–2 times longer than wide; teeth usually (6–)10–35 per side __ 2. L. horrida
21. Plants erect, low and stiffly, densely branched; leaf-blades narrowly triangular to narrowly elliptic, mostly 2–3 times longer than wide; teeth usually 3–6 per side ______3. L. leonardiorum