It does not seem particularly prudent to
try to make inference about bird density in a region midway
between your sites, that is at least a hundred kilometers from
any of your sampling effort. Seems that would stretch your
inference beyond the tolerance limits. Analysing the sites
separately would be the more statistically sound approach to
take. That being said, you'll also want to ask whether
relationships you find from along roadsides will remain relevant
throughout the site to which you wish to make inference.
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