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Mariana Doering, Maurício S. Garcia, Jeung Hee Schiefelbein, Leonardo Kerber & Rodrigo T. Müller (2025)
New tetrapod remains help constrain the age of a peculiar assemblage, including early dinosaurs and pterosaur precursors, from the Upper Triassic of southern Brazil
Journal of Vertebrate Paleontology e2552552
doi:
https://doi.org/10.1080/02724634.2025.2552552 https://www.tandfonline.com/doi/full/10.1080/02724634.2025.2552552The Hyperodapedon Assemblage Zone (AZ) is a key global marker of the Carnian stage in the Late Triassic. In the Candelária Sequence of Brazil (Rio Grande do Sul), it splits into the lower Hyperodapedon Acme Zone and the upper Exaeretodon Sub-AZ. Here we report new tetrapods from the previously overlooked lower exposures of the Niemeyer Complex (Agudo municipality). Intensive fieldwork yielded dozens of new specimens, including multiple hyperodapedontine rhynchosaur elements, as well as a partial skeleton referable to “Hyperodapedon” sanjuanensis via anatomical comparison and phylogenetic analysis, a lagerpetid, an early-diverging sauropodomorph, and a cynodont. This discovery provides evidence for the first definitive record of the Hyperodapedon Acme Zone in Agudo, with its fauna closely matching other late Carnian sites of southern Brazil, such as Buriol and Cerro da Alemoa, as well as correlative assemblages across southern Pangea. In contrast, the upper Niemeyer fauna is considered to be younger (latest Carnian–earliest Norian transition or early Norian) in age. Distinct lithologies, contrasting fossil assemblages, and the presence of index taxa support recognizing the lower and upper Niemeyer exposures as separate biozones. Our findings refine intercontinental correlations of Triassic tetrapods, enhance understanding of hyperodapedontine postcranial anatomy, and constrain the age of the lower beds of the Niemeyer Complex to the late Carnian.
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