Yet another thought, and please let me know if I'm off.
What I find intriguing is the progressive nature of the issue. Ordinarily, I would expect things to either go from too little infill to perhaps eventually normalizing or to start out squashed and eventually normalizing. Here's my personal experience:
- When I initially had my head calibrated, it was too close to the bed. First layers were "squashed" and the head was dragging over the PLA. After several layers, the print got better, eventually turning out good layers one after another.
- When I re-calibrated, I got the head a little too far. This resulted in the first layer looking a bit "loose" (but not quite "under extruded") with each subsequent layer getting better until it just looked good overall.
- Finally, when I got the first layer perfect, all layers looked uniform.
What I think is interesting is the layers don't follow any of these patterns. It seems initially like my second scenario - first layer too loose with things filling in on subsequent layers. However, instead of normalizing so that layers eventually turn out uniform, it seems like they get progressively more squashed.
Could it be that this is a two-fold issue: first layer is "loose" due to the head being just a little too high followed by the Z movements being just a little too small, meaning layers get progressively more squashed?
Let me know if my explanation doesn't make sense.