Dear Anne,
There are constant characters in the file you provide. That means that you can not use the Mkv model which assumes there are no constant characters in the matrix.
If you want to keep the constant characters and use Mk model then you also need to specify in CHARSTATELABELS how many possible states a character can take. BEAUti automatically partitions the matrix in groups of characters having the same number of possible states which is either taken from CHARSTATELABELS or calculated from the character. Character that can only take one state are not allowed as they don't bring any information. Note, that it is different from having a constant character which could take more than one state. Then if there is no
CHARSTATELABELS and there is a constant character, BEAUti can not assign the number of possible states greater than one. You can see an example of CHARSTATELABELS in this file:
you can also look at this paper: