Hi Ryan,
It is possible to start with a 'root' to your graphical model that is not 1. This is the approach I used in the analysis of my field experiment (R code is available
here).
It's also possible to include a "sampling" stage where not all the units of observation at the previous stage make it to the next. For example, I used this when counting all the seeds in only a subsample of pods (my code
here. Charlie has a better example that I modified mine from in one of the tech reports):
# Set up data for aster modeling through seeds
# Subsampling seed pods to include seeds/pod in life history graph
# graph:
# 1 -> e.surv -> fecund -> seed.pods -> pods.counted(sample) -> any seeds -> seeds.counted
# samp is binomial with sample size pods and known success probability p
That said, since your germination treatment wasn't explicitly sampled with this analysis in mind, it might not work well.
In your case, you appear to have (at least?) two experiments:
A) Germination
B) Life history
so it seems you should just analyze them separately, using aster from the start point of an individual seedling that germinated, and a GLM for germination. Maybe not the answer you were hoping for...but probably easier!