Unenlagiidae phylogenetic relationships among Paraves

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Ben Creisler

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Aug 22, 2025, 11:30:22 AMAug 22
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Ben Creisler

A new paper:

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Matías J. Motta, Federico L. Agnolín, Federico Brissón Egli, Sebastián Rozadilla & Fernando E. Novas (2025)
Phylogenetic relationships of Unenlagiidae among Paraves (Dinosauria)
Journal of Systematic Palaeontology 23(1): 2529608
doi: https://doi.org/10.1080/14772019.2025.2529608
https://www.tandfonline.com/doi/full/10.1080/14772019.2025.2529608

In recent years, several studies on the osteology, myology and palaeobiology of southern paravians of the clade Unenlagiidae have considerably increased the information on the group. Nevertheless, their phylogenetic relationships in the theropod phylogenetic tree remain under discussion. Unenlagiids were proposed as early divergent members of Avialae or as members of Dromaeosauridae. Using as a base the most comprehensive known coelurosaurian data matrix (TWiG dataset), we aimed to test the phylogenetic affinities of unenlagiids. The dataset includes new available information, scorings based on direct observation of materials, and the addition of 15 taxa (including the Gondwanan Imperobator, Pamparaptor, Unenlagia paynemili and Overoraptor), and 23 characters. The phylogenetic trees obtained in the present analysis support the inclusion of all Gondwanan paravians in the clade Unenlagiidae, as early divergent members of Avialae, and outside of Dromaeosauridae. This emphasizes the importance of Unenlagiidae in interpreting the phylogenetic relationships and early evolution of avialans.

Tim Williams

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Aug 25, 2025, 4:25:00 AM (13 days ago) Aug 25
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According to the full tree (in Supplemental material) the Jinguofortisidae is resolved within Ornithothoraces, as the most basal clade of Enantiornithes.  So quite a step up - previously the jinguofortisids were found to be more basal than _Sapeornis_.   (Though this more crownward position might just be a consequence of the number of ornithothoracine taxa sampled.)

Curiously, the authors state that they "recovered microraptorians, unenlagiids and anchiornithids as birds".  But they also stick to the definition of Avialae as "all taxa closer to _Passer_ than _Troodon_ and _Dromaeosaurus_".  I assume Avialae = birds.  Since the authors also found Microraptoria to be a subclade of Dromaeosauridae, then microraptorians can't be members of Avialae - and so they therefore aren't birds.    



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Ben Creisler

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Aug 28, 2025, 12:28:32 PM (10 days ago) Aug 28
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The pdf is now free:

Free pdf:

Matías J. Motta, Federico L. Agnolín, Federico Brissón Egli, Sebastián Rozadilla & Fernando E. Novas (2025)
Phylogenetic relationships of Unenlagiidae among Paraves (Dinosauria)
Journal of Systematic Palaeontology 23(1): 2529608
doi: https://doi.org/10.1080/14772019.2025.2529608
https://www.tandfonline.com/doi/full/10.1080/14772019.2025.2529608
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