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to mod-pagespeed-discuss
I installed ModPagespeed on debian/apache from the .deb and am using the default Pagespeed config. I run a handful of Virtual Hosts, and while modpagespeed is running, it does not seem to minifying CSS.
Once PageSpeed minifies it, it should keep serving the optimized
version as long as your site keeps getting traffic. The cache
lifetime you have set on the input files is 300s, though, which means
that if you get fewer than one hit per 300s you will probably be
serving some traffic unoptimized. (If that's happening, you could
turn on ModPagespeedLoadFromFile or increase your origin cache
lifetime.)