Nice find indeed - thanks for posting the pictures Nick!
I don't think we should assume these are a development version unless there's some direct evidence of that status (like a stamp of ОПЫТНАЯ - experimental). Absent that, it's more likely that these are production versions, just using an earlier internal construction than the one we are more used to seeing. It's quite common for tubes to evolve internal construction over time without designating that in any change of the marking - those are usually tied to a particular electrical specification rather than fine physical detail.
It'd be interesting to see the glow colour - it wouldn't surprise me if they are neon only, lacking the blue haze that comes from mercury addition. Digit stacking order might be different too.
Don't like the apparent corrosion on the pins though - would worry whether this might lead to micro-cracks in the glass and potentially compromise the gas seal. Personally I'd be tempted to keep them as historical specimens rather than working clock tubes unless you've got a hatful of them.
Jon.