Tube matching should be pretty critical as the push-pull on the mylar should be as symmetric as possible (unlike "usual" ESL where it's less important). On the Bev it is important that the membrane travels equal distance in each direction when the same capacitance is formed/varied equally on both sides on either half of the sinus.
Life span? I would guess around 10-15 years (much depends how much you play them, how many times you turn them on/off). It may be a good idea to buy spares since who knows where tubes will be in the future... A bit of an investment but worthwhile I think. Also handy if a tube goes pair shaped suddenly.
I should add that I played myself with the original, experimental amp hand build and hand wired by Harold Beveridge (no printed circuits) in 2006. They were built in 1972 as a concept proof. It still worked perfectly with the original tubes and traffo as I connected it to a 2SW speaker! Some tube amp.
If the speakers haven't worked for a long long time, once you replace the tube, it might be a good idea to turn them on slowly via a variac in slow (25V, 35V, 50V, 75V, 90V, 115V, 10mins each), incremental steps. Check the traffo still ok. As you increase voltage, listen for any possible buzzing from the speaker(s). A variac on ebay should cost c.$100-150. Always good to have at home if you don't own one already.
I am guessing you haven't heard the speakers play yet. If you never heard a Beveridge play and you like/appreciate pure sound, you are in for a treat - some say shock.
AV.