On Mar 5, 6:23�pm, "Klaus Hellnick" <
khelln...@sbcglobal.net> wrote:
> "pnyikos" �wrote in message
>
> news:ba322d5c-ff51-4e98...@y4g2000yqa.googlegroups.com...
>
> "By the way, it seems that extant birds are also widely believed to
> have evolved from a single common post-KT ancestor. �Here is a reply
> to me back in 2000 by Harshman's old nemesis, Cal King:"
>
> Bullshit!!!
> Ducks, parrots, rattites and others already had diversified before the K-T
> extinction.
> This is what is widely believed.
And it is also widely believed that ungulates, insectivores,
Xenarthans, primates, and bats had done so as well.
Your point?
> You quote two well known crackpots, who contradict each other.
> Klaus
What are your credentials for making such a statement? Let's put them
to the test.
Let's see whether you can do better than Harshman in dealing with the
following post of mine from last year. His reply, to put it mildly,
was underwhelming.
On Apr 26, 6:41 pm, John Harshman <
jharsh...@pacbell.net> wrote:
> pnyikos wrote:
> > On Apr 26, 10:52 am, John Harshman <
jharsh...@pacbell.net> wrote:
> >> pnyikos wrote:
> >>> On Apr 24, 10:41 am, John Harshman <
jharsh...@pacbell.net> wrote:
> >>>> pnyikos wrote:
> >>>>> John Harshman wrote:
> >>>>>> And doubtless there would be some
> >>>>>> other term for anything more closely related to Morganucodon than to
> >>>>>> Marmota.
> >>>>> Not if ranks were abolished. How would you quantify "more closely
> >>>>> related"?
> >>>> Surely you know this if you just think about it.
>
> >>> I have known since childhood that the Linnean classification has a
> >>> way. Your suggestions below are flawed.
>
> >> You have known wrong. Arbitrary dividing lines don't do a good job even
> >> of quantifying similarity, much less relationship.
>
> > The word "arbitrary" is extremely ambiguous. Put it this way: the
> > fewer and more insignificant the apomorphies, the smaller the Linnean
> > taxon. Thus the hypothetical young digger could be quite
> > communicative ("within error bars" to use an expression familiar to
> > you) if he said, "these morganucodonts are in the same subfamily as
> > Morganucodon, whereas the examples YOU name aren't even in the same
> > infraclass."
>
> I've asked you several times for the criteria you use to assign ranks to
> taxa. No response so far. How few is few? How insignificant is
> insignificant? How small is small?
I'm sure the systematists of old had a very good feel for this kind of
thing--just from a lifelong interest in the various Linnean taxa.
It's obvious, for instance, that the shoulder girdle of the platypus
should weigh far, far more in the classification than a dozen or so
mitochondrial genes.
These were presumably what led one article to classify it in a clade
with marsupials that excluded placentals. We've talked about this
before, and you agreed we shouldn't put too much faith in such
analyses.
Where the shoulder girdle is concerned, I'm not just talking about the
disappearance in all marsupials and placentals of the interclavicle
and procoracoid, primitive features shared by the platypus,
pelycosaurs, and therapsids.
One could write that off as homoplasy. But on p. 279 of Romer's
classic _Vertebrate Paleontology_, there is a picture of four shoulder
girdles, and the scapula of the "Virginia opossum" *Didelphis* has a
shape and mid-ridge that is very much like that of all placentals that
I have ever seen, while the one for the platypus is much more similar
to that of the pelycosaur and the therapsid that is in the
reproduction.
That feature, all by itself, cries out for putting marsupials and
placentals in a clade that excludes the platypus, and it would take a
huge number of countervailing characters to overturn that assessment,
unless they happened to be as striking as the shoulder girdle
differences.
> >>>> "Sharing a more recent
> >>>> common ancestor"
> >>> That might backfire. The way the tree of life is rooted, Morganucodon
> >>> might not have a common ancestor with anything except the huge clade
> >>> in which Mormota is located, of which the genus Morganucodon is thus
> >>> the sister group.
> >> I am unable to interpret that confusing statement. Everything has a
> >> common ancestor with everything else.
>
> > Sorry, I should have added "more recent" between "might not have a"
> > and "common ancestor". It was late and I was getting sleepy.
>
> Still doesn't make sense.
Morganucodon had various common ancestors with everything else [except
maybe prokaryotes] but none of them might have been more recent than
the one it shared with all of Mammalia and most of "Mammaliforma".
Does this still not make sense to you?
[snip side issue which is sure to come up later, on a more appropriate
thread]
> >>> If what I said above is true, and Morganucodon had no descendants, the
> >>> least inclusive clade containing Morganucodon besides *Morganucodon*
> >>> itself, is the huge clade containing Eutheria, and probably
> >>> Metatheria.
> >> Better than that. Morganucodon is actually outside Mammalia as commonly
> >> defined these days (i.e. as a crown group). It's a mammaliaform. How is
> >> this a problem?
>
> > The concept of "mammaliform" could be a problem if the vexing question
> > of where monotremes belong takes yet another turn. Because then a lot
> > of "mammaliform" creatures -- essentially ALL known Jurassic mammals
> > -- could suddenly become part of Mammalia.
>
> Not a problem for me. Is it a problem for you?
No, because I keep carrying old paraphyletic groups around in my head,
and this only requires a minor adjustment for me.
> > In Romer's day, ALL Jurassic mammals then known were widely believed
> > to be in the crown group formed by the extant mammals. Romer himself
> > thought that there was good evidence that all known Jurassic mammals
> > were "more advanced" than monotremes, and hence "more closely related"
> > cladistically speaking, to placentals and marsupials.
>
> > Item: the molars of the Cretaceous monotreme *Steropodon* and even the
> > Miocene (or is it Pliocene? I forget.) platypus *Obdurodon* look more
> > like the teeth of the triconodont *Priacodon* than the molars of a
> > "pantothere" or a "symmetrodont" or those of a Metatherian or
> > Eutherian.
> > So, unless the countervailing evidence is strong, all of these
> > critters could become part of "Mammalia" from which (if Mammalia is a
> > crown group, as you say) they are currently excluded.
>
> Not a problem. Even if "look more like" is counted as a synapomorphy.
>
> > On the other hand, Morganucodon would probably continue to be
> > excluded, because the quadrate and articular bones have not yet become
> > the malleus and incus as they have in monotremes. This key change is
> > IMO the best way to define "Mammalia".
But -- stop the presses! Today I saw in Wikipedia that there is a
(controversial) theory that this changeover is a case of convergent
evolution between monotremes and other extant mammals. The references
are:
Rich, T. H.; Hopson, J. A.; Musser, A. M.; Flannery, T. F.; & Vickers-
Rich, P. (2005). "Independent origins of middle ear bones in
monotremes and therians.". Science (Science) 307 (5711): 910 -
914. doi:10.1126/science.1105717. PMID 15705848. 10.1126/science.
1105717.
^ "Comment on "Independent Origins of Middle Ear Bones in Monotremes
and Therians" (I)". Science Magazine. Retrieved 2007-10-21.
^ "Comment on "Independent Origins of Middle Ear Bones in Monotremes
and Therians" (II)". Science Magazine. Retrieved 2007-10-21.
When I get into the University, I'll see just how strong the _Science_
paywall is; if I can't see these items, I'll take a trek down to the
library, but perhaps not today.
If this theory triumphs, then the dividing line between traditional
Reptilia and Mammalia will have to be different from the one I was
using, because polyphyletic taxa were shunned by Linnean
paleontologists as much as they are by people who use
"phylogenetic" [read: cladistic] classifications.
Peter Nyikos