I fail to see how there can be modules if they are not independent. To
me, the entire organism becomes a module.
I agree that for the purpose of study we need to define our unit
interest. But we still need to be informed on the developmental links
among body parts before defining our study region. If you wanted to
study the size of the vertebrate eye and relate it to being nocturnal
or diurnal, you would not want to study the size of the eyeball
separately from the size of the eye socket. In my opinion, you would
not have defined a module. If you were to study the eye socket and eye
size then you would have a more appropriate working definition.