Carleton Evolution and Palaeontology

I know this is way after the fact, but I was thinking of resurrecting some posts on the board in

I fail to see how there can be modules if they are not independent. To me, the entire organism

This paper may also be relevant here: http://sysbio.oxfordjournals.org/content/56/6/879.full.pdf+html

Looking at their ancestral state reconstruction it would appear that TSD is derived from ancestors